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In densely forested habitats, the territory controlled by a given chimpanzee group is only a few kilometers wide. In sparsely wooded regions, it can be as much as thirty kilometers across. These are the territories that a chimp group considers its turf, its home, its fatherland or motherland, to which something like patriotic sentiments are owed. It is not to be trespassed by strangers. It’s a jungle out there. The typical day range of a chimp combat patrol is a few kilometers. So if they live in heavy forest, they can fairly readily patrol a good portion of the border in a single day. But if the vegetation and food supply are more sparse and their territory accordingly larger, it may be a few days’ journey from one end to the other, and longer if they go around the perimeter.A patrol is typified by cautious, silent travel during which the members of the party tend to move in a compact group. There are many pauses as the chimpanzees gaze around and listen. Sometimes they climb tall trees and sit quietly for an hour or more, gazing out over the “unsafe” area of a neighboring community. They are very tense and at a sudden sound (a twig cracking in the undergrowth or the rustling of leaves) may grin and reach out to touch or embrace one another.During a patrol the males, and occasionally a female, may sniff the ground, treetrunks, or other vegetation. They may pick up and smell leaves, and pay particular attention to discarded food wadges, feces, or abandoned tools on termite heaps. If a fairly fresh sleeping nest is seen, one or more of the adult males may climb up to inspect it and then display around it so that the branches are pulled apart and it is partially or totally destroyed.Perhaps the most striking aspect of patrolling behavior is the silence of those taking part. They avoid treading on dry leaves and rustling the vegetation. On one occasion vocal silence was maintained for more than three hours … [When] patrolling chimpanzees return once more to familiar areas, there is often an outburst of loud calling, drumming displays, hurling of rocks, and even some chasing and mild aggression between individuals … Possibly this noisy and vigorous behavior serves as an outlet for the suppressed tension and social excitement engendered by journeying silently into unsafe areas.31


In this description by Jane Goodall of a patrol at Gombe, we are taken by the ability of the chimps to overcome their fear, to exercise self-control by inhibiting their usual noisy interchanges, but particularly by their deductive abilities. These chimps are tracking. They are weighing the evidence of branches, footprints, droppings, artifacts. We might expect that, when food is in short supply, group differences in tracking skills help determine who lives and who dies. Not just strength and aggressiveness are being selected here, but something akin to reasoning and quick-wittedness. And stealth. When one human who lived with a troop for a long time tried to accompany a patrol as it set out, they looked at him reprovingly. He was just too clumsy. He could not, as they do, slip silently through the forest.

So the long-range combat patrol wends its way toward the borders of their turf If it’s more than a day’s walk, they’ll set up camp at night and continue on their way tomorrow. What happens if they encounter members of another group, Strangers from the adjacent territory? If it’s just one or two intruders, they’ll attempt to attack and kill them. There’s much less disposition here toward threat displays and intimidation. But if two parties of roughly equal strength encounter one another, now there are a great many threat displays, rocks and sticks are thrown, trees are drummed. “Somebody hold me back, I’m gonna break his knees,” you can almost hear them saying. They practice threat assessment If the patrol senses an obviously larger number of Strangers, it is likely to beat a hasty retreat. At other times chimp patrols may penetrate enemy territory or even raid its populated core area—for many purposes, including copulating with unfamiliar females. The combination of tracking, stealth, danger, teamwork, fighting hated enemies, and the opportunity for sex with strange females is enormously attractive to the males.

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